Deforestation and forest degradation are causing widespread loss of tropical biodiversity, profoundly impacting ecosystem functioning as well as stocks of natural resources and ecosystem assets (natural capital). The greatest reductions in diversity are experienced as forests are converted to permanent agriculture, a process that disrupts the delivery of important ecosystem services such as pollination and pest control. In contrast, the impacts from well-managed smallholder agriculture are less extreme, as the associated land parcels are typically embedded within landscape mosaics comprising fallows and forest remnants. Wallacea is currently emerging as a new developmental frontier in Indonesia and a target for agribusiness and extractive industries. A particularly understudied part of the Asian tropics, it has an exceptionally distinctive vertebrate diversity which forms the second highest level of endemism in the world, making the region a global priority for both conservation and ecosystem service provision. In addition, land-use history and current trajectories remain poorly understood, with the region notably omitted from recent deforestation baselines for this very reason. In fact, the diverse history of the Wallacea archipelago is acknowledged as a major source of uncertainty when applying land-use change models developed from elsewhere in Southeast Asia, as well as predicting the impacts of future environmental change. Given that further forest degradation and agricultural conversion are expected in Wallacea, the future prospects for natural capital in the region depend to a large extent on how we manage human-modified landscapes. Bringing together an interdisciplinary team from British and Indonesian universities, with four NGO partners active in Wallacea, this project will deliver the science needed to understand tensions in land-use and the responses of biodiversity to environmental change in Wallacea. We propose a novel and ambitious study of biodiversity responses to recent and historical land-cover change across multiple landscapes in little-studied islands, so that evaluations of current land-use policies and predictions of future environmental scenarios will be evidence-based and realistic. More specifically, spatial trajectories of land-cover change will be generated for each landscape, drawing from publicly-available remote-sensed data and local land-use plans. This will enable us to hindcast forest cover back to Wallace's time and forecast to key target years for international policy commitments (e.g. 2030 for the UN Sustainable Development Goals and 2050s for the UN Framework on Climate Change). Significantly, we will generate new biodiversity data from across land-cover gradients in forests, agroforests and intensive farmland (e.g. cocoa, oil palm, coffee), model community responses to past, present and future forest cover, and apply state-of-the-art genomics methods to assess genetic and evolutionary responses to land-cover change for several important conservation flagship species (NERC-Ristekdikti programme's Goals 1 and 2). Focusing on terrestrial vertebrates, the fauna that environmental policies aiming at safeguarding biodiversity are typically focused upon, we will track Alfred Russel Wallace's journey through Sulawesi and the Moluccas (Maluku). Finally, with collaboration from Project Partners with additional expertise in Wallacea, we will evaluate the impact of current land-use policies on ecosystem assets and dependent human beneficiaries, drawing on our land-cover, biodiversity and additional carbon and socio-economic data (Programme's Goals 2 and 3). Because many provinces in Indonesia are yet to implement newly-required spatial planning processes, our joint environmental research has an unprecedented opportunity to inform local development.

This project proposes to form a consortium of partners from the United Kingdom, Tanzania, Rwanda, Bolivia, Brazil and the United States to develop a research framework that will help fill knowledge gaps related to how climate change will impact provisioning and regulatory ecosystem services; how these changes might affect rural livelihoods; and how governance solutions can be developed to help manage those changes in countries of the Amazon and Sub-Saharan Africa. This research will generate new data and understandings while building developing country capacity to design and implement policy relevant research on the impacts of climate change on ecosystem services and rural livelihoods. Our research will be conducted at four sites where the ecosystem services provided by forests and hydrological systems are highly interdependent ecologically and integral to local and, in most cases, national economies, and are at risk of major disruption from climate change. These sites include the Great Ruaha river landscape in Tanzania; the Nyungwe forest in Rwanda; the Madidi landscape of Bolivia; and the flooded (várzea) forest landscape in Mamiraua, Brazil. We will develop systems dynamic models of climate, ecosystem services and rural livelihoods in these landscapes, and combine these with participatory assessments of governance options, across a range of spatial and political scales. The models will acknowledge that regulatory and provisioning services are ecologically interdependent, which will permit an exploration of synergies and trade-offs in these ecosystem services under various management regimes. The role that hydrological systems and watershed management play in regulating flows of provisioning services is the common thread that binds these sites, however, the ecological and socio-political differences across sites will make the policy implications of research results broadly applicable to other locations in the Amazon and Sub-Saharan Africa. Combined, the sites will allow us to speak to a variety of water-livelihood interactions related to fisheries, energy, agriculture, sanitation and tourism, and to a range of governance contexts. These are sites where the consortium's existing data bases and local relationships are especially well developed, allowing us to conduct both quantitative modeling and qualitative research with the greatest effectiveness and efficiency. Through a series of informal preparatory meetings, consortium members and local partners will initiate a participatory process to design a research program to assess the impact of climate change on ecosystem services provision and local livelihoods. Through WebEx sessions, we will host virtual meetings to exchange ideas that will lay the foundation for a consortium planning workshop in Bolivia to consolidate the interdisciplinary team and identify research needs at each site. The workshop will produce a research program designed to identify major knowledge gaps in existing data and capacity building priorities and estimate the level of funding required to support the proposed research. This grant will enable us to develop a framework for building a cadre of researchers with the skills needed to assess climate change impacts on ecosystem services and rural livelihoods for informing policy makers. By working collaboratively across sites, the program will foster direct south-south exchange of skills and knowledge and build the collegial relationships needed for future joint research. Strengthening capacity to conduct policy relevant research is critical for guiding development decisions that enhance local and national resilience to ecological, economic and social shocks linked to climate change. As a result of this proposed program, our partners and their communities in developing nations will be better prepared to adapt to climate change and to manage ecosystem services for the benefit of the rural poor.